The king was a gorilla called Titus. Although he had been deposed by his son, death seemed to have restored him to his full glory. The mighty silverback was once the dominant head of a tight-knit group of the great apes whose kingdom was on the eastern slopes of the Visoke volcano in Rwanda's border lands, and a vital figure in the battle for the survival of the species.
Titus, the most studied and photographed ape in history, has died
