Populations of some of the world's largest wild animals are dwindling, raising the threat of an "empty landscape", say scientists.
About 60% of giant herbivores - plant-eaters - including rhinos, elephants and gorillas, are at risk of extinction, according to research.
Analysis of 74 herbivore species, published in Science Advances, blamed poaching and habitat loss.
A previous study of large carnivores showed similar declines.
Prof William Ripple, of Oregon State University, led the research looking at herbivores weighing over 100kg, from the reindeer up to the African elephant.
"This is the first time anyone has analysed all of these species as a whole," he said.
"The process of declining animals is causing an empty landscape in the forest, savannah, grasslands and desert."